Balanomorph cirripedes from the Eocene–Oligocene of the Hampshire Basin (United Kingdom) and the Middle Eocene of the Cotentin Peninsula, Manche (France) are described. A new genus, Vectibalanus, is founded, with the type species Balanus unguiformis J. de C. Sowerby, 1846; assigned to this are also Balanus erisma J. de C. Sowerby, 1846 and Vectibalanus mortoni sp. nov. In addition, a new species of Lophobalanus Zullo, 1984, L. fresvillensis sp. nov., is described. This is the first record of that genus from outside the eastern USA and the oldest species known to date. Cladistic analysis of 24 morphological characters suggests that Vectibalanus unguiformis is sister taxon to a group comprising the most derived balanomorph taxa, and thus represents an important transition in the evolution of the group, with the initiation of development of a complex parietal wall structure. Vectibalanus unguiformis was evidently adapted to low salinity habitats (10–30 ppt), and is the oldest known brackish water barnacle. The other species ( V. erisma, V. mortoni sp. nov.) occupied more clearly marine environments (>30 ppt). Balanomorph barnacles appeared simultaneously in the Priabonian (Upper Eocene) of the Gulf and Atlantic seaboards of the USA and northwest Europe, which probably represents a northerly migration from Tethys.

Successions exposed in the Agadir Basin (upper Albian to middle Turonian), in the Anti-Atlas (lower Turonian) in Morocco and in central Tunisia (Cenomanian–Turonian) yield abundant microcrinoids of the family Roveacrinidae, which are described and assigned to 32 species and formae, in ten genera. The following new taxa are described: Fenestracrinus gen. nov. with the type species F. oculifer sp. nov., Discocrinus africanus sp. nov., Styracocrinus rimafera sp. nov., Lebenharticrinus quinvigintensis sp. nov., L. zitti sp. nov., Euglyphocrinus cristagalli sp. nov., E. jacobsae sp. nov., E. truncatus sp. nov., E. worthensis sp. nov., Roveacrinus gladius sp. nov., R. solisoccasum sp. nov. and Drepanocrinus wardorum sp. nov. In addition, the new subfamily Plotocrininae is erected. The stratigraphical distribution of the taxa in two important localities, Taghazout in the Agadir Basin (Morocco) and Sif el Tella, Djebel Mhrila (central Tunisia), is provided. The faunas from the uppermost Albian and lowermost Cenomanian of the Agadir Basin are nearly identical to those recorded from central Texas, USA, some 5,300 km away, and permit a detailed correlation (microcrinoid biozones CeR1 and CeR2) to be established across the southern part of the Western Tethys, independently supported by new ammonite records. For the middle and upper Cenomanian, rather few detailed records of microcrinoids are available elsewhere, and the North African record forms the basis for a new zonation (CeR3–CeR6). The distribution of Turonian Roveacrinidae in North Africa is evidently very similar to that described in the Anglo-Paris Basin, and zones TuR1–3, TuR9, 10 and 14 are recognised for the first time in the Tethys.